Protei 5.3 Surface control was really a preparation for what would become Protei 6.0 "Batman".
The spines will connected the ribs through the holes. And different motors will be used to drive different segments, which will make each segments totally controllable. But with different motors at different places, might be a little complicated. Another issue is that since the spines are flexible, and there is only one central spine, what will happen is that the hull may twist when bending and this twist has to be avoided.
Prototype prepared by